Publications

Sarath R Nair, Adrián Palacios-Muñoz, Sage Martineau, Malak Nasr, Jan Clemens

Sex-specific behavioral feedback modulates sensorimotor processing and drives flexible social behavior

2026, accepted at Nature Communications

• bioRxiv preprint
• Code

Stefan Dahlhoff, Sander Liessem, Fathima Mukthar Iqbal, Adrián Palacios-Muñoz, Federico Cascino-Milani, Mert Erginkaya, Aleyna M Diniz, E Axel Gorostiza, Ansgar Büschges, Jan Clemens, and Jan M Ache

Control of walking direction by descending and dopaminergic neurons in Drosophila

2025, in revision

• bioRxiv preprint
• Code

Luiza F Passos, Anja Hutschenreiter, Margarita B Jaramillo, Emanuele Cissello, Jan Clemens, Filippo Aureli, Serge Wich

Breaking through the noise in passive acoustic monitoring: Evaluating commercial and open-source tools for automated detection of highly variable primate vocalizations

2025, submitted

• Code

Adrián Palacios-Muñoz, Jan Clemens

Drosophila males integrate song and pheromones using context-specific strategies

2025, in revision

• PDF
• Code

Giacomo Costalunga, Carolina C Sanchez, Jonathan I Benichov, Colin Cross, Jan Clemens, Daniela Vallentin

Attuning to song duels facilitates song-matching in nightingales

2025, submitted

• bioRxiv preprint
• Code

Adrienne Chitayat, Jan Clemens, Catherine Crockford, Anne-Sophie Crunchant, Ammie K Kalan, Alex Piel, Fiona Stewart, Serge Wich

Acoustics, balance, and chimpanzees – The ABCs of developing a deep learning-based automated acoustic detector for wild chimpanzee (Pan troglodytes) loud calls

2025, under review

• Preprint
• Code

Winston Mann, Bettina Erregger, Matthias Hennig, Jan Clemens

Resonant song recognition and the evolution of acoustic communication in crickets

2025, iScience

• PDF
• bioRxiv preprint
• Code

Elsa Steinfath, Afshin Khalili (co-first author), Melanie Stenger, Bjarne L Schultze, Sarath N Ravindran, Kimia Alizadeh, Jan Clemens

A neural circuit for context-dependent multimodal signaling in Drosophila

2025, Nature Communications

• PDF
• bioRxiv preprint
• Code

Ava Kiai, Jan Clemens, Manfred Kössl, David Poeppel, Julio C. Hechavarría

Flexible control of vocal timing in Carollia perspicillata bats enables escape from acoustic interference

2023, Communications Biology

• PDF
• Code

Roshan Kumar Vijendravarma, Sunitha Narasimha, Elsa Steinfath, Jan Clemens, Pierre Leopold

Drosophila females have an acoustic preference for symmetric males

2022, PNAS, 119(13):e2116136119

• PDF
• Code

Jan Clemens, Mala Murthy

Quadratic and adaptive computations yield an efficient representation of song in Drosophila auditory receptor neurons

2021, in review

• bioRxiv preprint
• PDF
• Code and data
• More code

Jan Clemens, Stefan Schöneich, Konstantinos Kostarakos, Matthias Hennig, Berthold Hedwig

A small, computationally flexible network produces the phenotypic diversity of song recognition in crickets

2021, eLife, 10:e61475

• PDF
• bioRxiv preprint
• Code

Elsa Steinfath, Adrian Palacios, Julian Rottschäfer, Deniz Yuezak, Jan Clemens

Fast and accurate annotation of acoustic signals with deep neural networks

2021, eLife, 10:e68837

• PDF
• bioRxiv preprint
• Documentation and code

Jan Clemens, Bernhard Ronacher, Michael Reichert

Sex-specific speed-accuracy tradeoffs shape neural processing of acoustic signals in a grasshopper

2021, Proceedings of the Royal Society B, 287:20210005

• PDF
• bioRxiv preprint
• Code

Deutsch, D, Pacheco, DA, Encarnacion-Rivera, L, Pereira, T, Fathy, Clemens, J, Girardin, C, R, Calhoun, A, Ireland, EC, Burke, AT, Dorkenwald, S, McKellar, C, Macrina, T, Lu, R, Lee, K, Kemnitz, N, Ih, D, Castro, M, Halageri, A, Jordan, C, Silversmith, W, Wu, J, Seung, HS, and Murthy, M.

The neural basis for a persistent internal state in Drosophila females.

2020, eLife, 9:e59502

• PDF

Jan Clemens, Matthias Hennig

Coding strategies in insects

2020, The Senses - Audition (2nd edition), book chapter

• PDF

David Deutsch, Jan Clemens (co-first author), Stephan Thiberge, Georgia Guan, and Mala Murthy

Shared song object detector neurons in Drosophila male and female brains drive divergent, sex-specific behaviors

2019, Current Biology, 23:3200-3215

• PDF
• bioRxiv preprint
• Code

Christa Baker, Jan Clemens (co-corresponding author), and Mala Murthy

Acoustic pattern recognition and courtship songs: insights from insects

2019, Annual Reviews of Neuroscience, 42:129-147

• PDF

Jan Clemens*, Philip Coen*, Frederic A. Roemschied*, Talmo Pereira, David Mazumder, Diego Pacheco, and Mala Murthy (* eq. contribution)

Discovery of a new song mode in Drosophila reveals hidden structure in the sensory and neural drivers of behavior

2018, Current Biology, 28:2400–2412

• PDF
• dispatch
• press
• bioRxiv preprint
• Code/data 1
• Code/data 2

Jan Clemens, Nofar Ozeri, Mala Murthy

Fast intensity adaptation enhances the encoding of sound in Drosophila

2018, Nature Communications, 9:134

• PDF
• bioRxiv preprint

David Stern, Jan Clemens, Pip Coen, Adam Calhoun, Troy Shirangi, John Hogenesch, Ben Arthur, Mala Murthy

Experimental and statistical reevaluation provides no evidence for Drosophila courtship song rhythms

2017, PNAS, 114(37):9978-9983

• PDF
• bioRxiv preprint
• Code&data

Jan Clemens, Jennifer Aufderheide (eq. contribution), Bernhard Ronacher

Relative weighting of acoustic information during mating decisions in grasshoppers indicates signatures of sexual selection

2017, Journal of Comparative Physiology A, 203(11):891-901

• PDF
• Code&data

Jan Clemens, Mala Murthy

The use of computational modeling to link sensory processing with behavior in Drosophila

2017, Celik, A., Wernet, M. (eds.) Decoding neural circuit structure and function

• Book chapter
• PeerJ preprint
• PDF

Philip Coen, Marjorie Xie, Jan Clemens, Mala Murthy

Sensorimotor transformations underlying variability in song intensity during Drosophila courtship

2016, Neuron, 89(3):629–644

• PDF
• preview

Jan Clemens, Cyrille Girardin (eq. contribution), Philip Coen, Georgia Guan, Barry Dickson, Mala Murthy

Connecting neural codes with behavior in the auditory system of Drosophila

2015, Neuron, 87(6):1332-1343

• PDF
• preview
• press

Florian Rau, Jan Clemens, Viktor Naumov, Matthias Hennig, Susanne Schreiber

Firing-rate resonances in the peripheral auditory system of the cricket Gryllus bimaculatus

2015, Journal of Comparative Physiology A, 201(11):1075-1090

• PDF

Jan Clemens, Florian Rau, Matthias Hennig, Jannis Hildebrandt

Context-dependent coding and gain control in the auditory system of crickets

2015, European Journal of Neuroscience, 42(7):2390-2406

• PDF

Kelly LaRue, Jan Clemens, Gordon Berman, Mala Murthy

Acoustic duetting in Drosophila virilis relies on the integration of auditory and tactile signals

2015, eLife, 4:e07277

• PDF

Jan Clemens, Stefanie Krämer (eq. contribution), Bernhard Ronacher

Asymmetrical integration of evidence during courtship decisions in grasshoppers

2014, PNAS, 111(46):16562–16567

• PDF
• press release

Bernhard Ronacher, Matthias Hennig, Jan Clemens

Computational principles underlying the recognition of acoustic signals in grasshoppers and crickets

2014, Journal of Comparative Physiology A, 201(1):61-71, invited review

• PDF

Matthias Hennig, Klaus-Gerhard Heller, Jan Clemens

Time and timing in the acoustic recognition system of crickets

2014, Frontiers in Physiology, 5:286, invited review

• PDF

Philip Coen, Jan Clemens, Andrew Weinstein, Diego Pacheco, Yi Deng, and Mala Murthy

Dynamic sensory cues shape song structure in Drosophila

2014, Nature, 507:233-237

• PDF
• news&views

Jan Clemens, Bernhard Ronacher

Feature extraction and combination underlying decision making during courtship in grasshoppers

2013, Journal of Neuroscience, 33(29):12136-12145

• PDF

Jan Clemens, Matthias Hennig

Computational principles underlying the recognition of acoustic signals in insects

2013, Journal of Computational Neuroscience, 35(1):75-85

• PDF

Jan Clemens, Sandra Wohlgemuth, Bernhard Ronacher

Nonlinear computations underlying temporal and population sparseness in the auditory system of the grasshopper

2012, Journal of Neuroscience, 32(29):10053-10062

• PDF

Jan Clemens, Olaf Kutzki, Bernhard Ronacher, Susanne Schreiber, Sandra Wohlgemuth

Efficient transformation of an auditory population code in a small sensory system

2011, PNAS, 108(33):13812-13817

• PDF
• press release
• media coverage

Jan Clemens, Gerroth Weschke, Astrid Vogel, Bernhard Ronacher

Intensity invariance properties of auditory neurons compared to the statistics of relevant natural signals in grasshoppers

2010, Journal of Comparative Physiology A, 196:285-297

• PDF